Who was Jesus referring to with these scriptures?

Jesus was speaking to the religious leaders of his day. As with everything in the Bible, there is a minor fulfillment and a major fulfillment. Everything occurs twice in the Bible. The injection of paganism into the realm of "christianity" in 325 C.E. makes all those pagan rituals that God hates into modern traditions that the religious leaders of today continue to promote. Any religion that practices rituals and holidays of pagan origin are just as guilty as those religious leaders in Jesus' day in that they "nullify the word of God for the sake of your tradition"